Horses were first domesticated in the Pontic-Caspian steppes, northern Caucasus

 By whom and where were modern horses first domesticated? When did they conquer the rest of the world? And how did they supplant the myriad of other types of horses that existed at that time? This long-standing archaeological mystery finally comes to an end thanks to a team of 162 scientists specialising in archaeology, palaeogenetics and linguistics.

A few years ago, Ludovic Orlando's team looked at the site of Botai, Central Asia, which had provided the oldest archaeological evidence of domestic horses. The DNA results, however, were not compliant: these 5500-year-old horses were not the ancestors of modern domestic horses1. Besides the steppes of Central Asia, all other presumed foci of domestication, such as Anatolia, Siberia and the Iberian Peninsula, had turned out to be false. “We knew that the time period between 4,000 to 6,000 years ago was critical but no smoking guns could ever be found” says CNRS research professor Orlando. The scientific team, therefore, decided to extend their study to the whole of Eurasia by analysing the genomes of 273 horses that lived between 50,000 and 200 years BC. This information was sequenced at the Centre for Anthropobiology and Genomics of Toulouse (CNRS/Université Toulouse III - Paul Sabatier) and Genoscope2 (CNRS/CEA/Université d’Évry) before being compared with the genomes of modern domestic horses.

This strategy paid off: although Eurasia was once populated by genetically distinct horse populations, a dramatic change had occurred between 2000 and 2200 BC. “That was a chance: the horses living in Anatolia, Europe, Central Asia and Siberia used to be genetically quite distinct” notes Dr Pablo Librado, first author of the study. Then, a single genetic profile, previously confined to the Pontic steppes (North Caucasus)3, began to spread beyond its native region, replacing all the wild horse populations from the Atlantic to Mongolia within a few centuries. “The genetic data also point to an explosive demography at the time, with no equivalent in the last 100,000 years” adds Pr Orlando. “This is when we took control over the reproduction of the animal and produced them in astronomic numbers.”

But how can this rapid population growth be explained? Interestingly, scientists found two striking differences between the genome of this horse and those of the populations it replaced: one is linked to a more docile behaviour and the second indicates a stronger backbone. The researchers suggest that these characteristics ensured the animals’ success at a time when horse travel was becoming “global”.

The study also reveals that the horse spread throughout Asia at the same time as spoke-wheeled chariots and Indo-Iranian languages. However, the migrations of Indo-European populations, from the steppes to Europe during the third millennium BC4 could not have been based on the horse, as its domestication and diffusion came later. This demonstrates the importance of incorporating the history of animals when studying human migrations and encounters between cultures.

This study was directed by the the Centre for Anthropobiology and Genomics of Toulouse (CNRS/ Université Toulouse III – Paul Sabatier) with help from Genoscope (CNRS/CEA/Université d’Évry). The French laboratories Archéologies et sciences de l'Antiquité (CNRS/Université Paris 1 Panthéon Sorbonne/Université Paris Nanterre/Ministère de la Culture), De la Préhistoire à l'actuel : culture, environnement et anthropologie (CNRS/Université de Bordeaux/Ministère de la Culture) and Archéozoologie, archéobotanique : sociétés, pratiques et environnements (CNRS/MNHN) also contributed, as did 114 other research institutions throughout the world. The study was primarily funded by the European Research Council (Pegasus project) and France Genomique (Bucéphale project).

Some previous results of the Pegasus project:

• Scythian horses shed light on animal domestication, 27 April 2017;
• Unsaddling old theory on origin of horses, 22 February 2018.

Humans did not cause woolly mammoths to go extinct -- climate change did

 For five million years, woolly mammoths roamed the earth until they vanished for good nearly 4,000 years ago – and scientists have finally proved why.

The hairy cousins of today’s elephants lived alongside early humans and were a regular staple of their diet – their skeletons were used to build shelters, harpoons were carved from their giant tusks, artwork featuring them is daubed on cave walls, and 30,000 years ago, the oldest known musical instrument, a flute, was made out of a mammoth bone.

Now the hotly debated question about why mammoths went extinct has been answered - geneticists analysed ancient environmental DNA and proved it was because when the icebergs melted, it became far too wet for the giant animals to survive because their food source – vegetation – was practically wiped out.

The 10-year research project, published in Nature today (20 October 2021), was led by Professor Eske Willerslev, a Fellow of St John’s College, University of Cambridge, and director of The Lundbeck Foundation GeoGenetics Centre, University of Copenhagen.

The team used DNA shotgun sequencing to analyse environmental plant and animal remains – including urine, faeces and skin cells – taken from soil samples painstakingly collected over a period of 20 years from sites in the Arctic where mammoth remains were found. The advanced new technology means scientists no longer have to rely on DNA samples from bones or teeth to gather enough genetic material to recreate a profile of ancient DNA. The same technique has been used during the pandemic to test the sewage of human populations to detect, track and analyse Covid-19.

Professor Willerslev said: “Scientists have argued for 100 years about why mammoths went extinct. Humans have been blamed because the animals had survived for millions of years without climate change killing them off before, but when they lived alongside humans they didn’t last long and we were accused of hunting them to death.

“We have finally been able to prove was that it was not just the climate changing that was the problem, but the speed of it that was the final nail in the coffin – they were not able to adapt quickly enough when the landscape dramatically transformed and their food became scarce.

“As the climate warmed up, trees and wetland plants took over and replaced the mammoth’s grassland habitats. And we should remember that there were a lot of animals around that were easier to hunt than a giant woolly mammoth – they could grow to the height of a double decker bus!”

The woolly mammoth and its ancestors lived on earth for five million years and the huge beasts evolved and weathered several Ice Ages. During this period, herds of mammoths, reindeer and woolly rhinoceroses thrived in the cold and snowy conditions.

Despite the cold, a lot of vegetation grew to keep the various species of animals alive – grass, flowers, plants, and small shrubs would all have been eaten by the vegetarian mammoths who probably their tusks to shovel snow aside and are likely to have used their trunks to uproot tough grasses. They were so big because they needed huge stomachs to digest the grass.

Mammoths could travel a distance equivalent of going around the world twice during their lifetime and fossil records show they lived on all continents except Australia and South America. Populations were known to have initially survived the end of the last Ice Age in small pockets off the coasts of Siberia and Alaska – on Wrangel Island and St Paul Island – but the research found they actually lived longer elsewhere too and the breeds of mammoths on both the islands were closely related despite being geographically separated. As part of the project, the team also sequenced the DNA of 1,500 Arctic plants for the very first time to be able to draw these globally significant conclusions.

Dr Yucheng Wang, first author of the paper and a Research Associate at the Department of Zoology, University of Cambridge, said: “The most recent Ice Age – called the Pleistocene – ended 12,000 years ago when the glaciers began to melt and the roaming range of the herds of mammoths decreased. It was thought that mammoths began to go extinct then but we also found they actually survived beyond the Ice Age all in different regions of the Arctic and into the Holocene – the time that we are currently living in ­– far longer than scientists realised.

“We zoomed into the intricate detail of the environmental DNA and mapped out the population spread of these mammals and show how it becomes smaller and smaller and their genetic diversity gets smaller and smaller too, which made it even harder for them to survive.

“When the climate got wetter and the ice began to melt it led to the formation of lakes, rivers, and marshes. The ecosystem changed and the biomass of the vegetation reduced and would not have been able to sustain the herds of mammoths. We have shown that climate change, specifically precipitation, directly drives the change in the vegetation – humans had no impact on them at all based on our models.”

Humans lived alongside woolly mammoths for at least 2,000 years – they were even around when the pyramids were being built. Their disappearance is the last big naturally occurring extinction story. Our fascination with the huge beasts continues today with ‘Manny’ the woolly mammoth starring as the main character in five Ice Age animated films, and scientists hoping to resurrect them from the dead.

Professor Willerslev said: “This is a stark lesson from history and shows how unpredictable climate change is – once something is lost, there is no going back. Precipitation was the cause of the extinction of woolly mammoths through the changes to plants. The change happened so quickly that they could not adapt and evolve to survive.

“It shows nothing is guaranteed when it comes to the impact of dramatic changes in the weather. The early humans would have seen the world change beyond all recognition – that could easily happen again and we cannot take for granted that we will even be around to witness it. The only thing we can predict with any certainty is that the change will be massive.”

Early modern human from Southeast Asia adapted to a rainforest environment

 Traditional assumptions have often seen tropical rainforests as a barrier to early Homo sapiens. However, growing proof shows that humans adapted to and lived in tropical rainforest habitats of Southeast Asia. Some researchers also suggest that, in the past, other human species, like Homo erectus and Homo floresiensis, became extinct because they could not adapt to this environment as our species did. However, we know very little about the ecological adaptation of fossil humans, including what they were eating.

Zinc isotopes reveal what kind of food was primarily eaten

In this study, researchers analysed the zinc stable isotope ratios from animal and human teeth from two sites in the Huà Pan Province of Laos: Tam Pà Ling and the nearby site of Nam Lot. "The site of Tam Pà Ling is particularly important for palaeoanthropology and archaeology of Southeast Asia because it holds the oldest and most abundant fossil record of our species in this region," explains Fabrice Demeter, researcher at the University of Copenhagen. However, there is little archaeological evidence, like stone tools, hearth features, plant remains, cut marks on bones, in Tam Pà Ling: only teeth and bones. This makes isotopic approaches the only way to gain insight into past dietary reliance.

Nitrogen isotope analysis, in particular, can help scientists learn if past humans were eating animals or plants. However, the collagen in bones and teeth needed to do these analyses is not easily conservable. In tropical regions like the one at Tam Pà Ling this problem is even more acute. "New methods -- such as zinc isotope analysis of enamel -- can now overcome these limitations and allow us to investigate teeth from regions and periods we could not study before," says study leader Thomas Tütken, professor at the Johannes Gutenberg University's Institute of Geosciences. "With zinc stable isotope ratios, we can now study Tam Pà Ling and learn what kind of food our earliest ancestors in this region were eating."

First study that reveals the whole diet of fossil humans from Southeast Asia

The fossil human studied in this research dates from the Late Pleistocene, more precisely from 46,000 to 63,000 years ago. With it, various mammals from both sites, including water buffaloes, rhinos, wild boars, deer, bears, orangutans, macaques, and leopards, were also analysed. All these different animals show various eating behaviours, making for an ideal background to determine what exactly humans were eating at the time. The more diverse the animal remains found at a particular site are, the more information the researchers can use to understand the diet of prehistoric humans.

When we compare the zinc isotope values from the fossil Homo sapiens of Tam Pà Ling to that of the animals, it strongly suggests that its diet contained both plants and animals. This omnivorous diet also differs from most nitrogen isotope data of humans in other regions of the world for that time period, where a meat-rich diet is almost consistently discerned. "Another kind of analysis performed in this study -- stable carbon isotopes analysis -- indicates that the food consumed came strictly from forested environments," says Élise Dufour, researcher at the National Natural History Museum of Paris. "The results are the oldest direct evidence for subsistence strategies for Late Pleistocene humans in tropical rainforests."

Researchers often associated our species with open environments, like savannahs or cold steppes. However, this study shows that early Homo sapiens could adapt to different environments. Together, the zinc and carbon isotope results may suggest a mix of specialized adaptations to tropical rainforests seen from other Southeast Asian archaeological sites. "It will be interesting, in the future, to compare our zinc isotope data with data from other prehistoric human species of Southeast Asia, like Homo erectus and Homo floresiensis, and see if we could understand better why they went extinct while our species survived," concludes first author Nicolas Bourgon, a researcher at the Max Planck Institute for Evolutionary Anthropology.

Only one in four Western Roman emperors died of natural causes

The Roman Empire was ruled by 175 men, from Augustus (63 BCE-19 CE) to Constantine XI (1405-53), including the Eastern or Byzantine Empire after the split in 395 CE, but excluding those who did not rule in their own right because they were minors during regencies or co-emperors.

Only 24.8% of the 69 rulers of the Western Empire died of natural causes. The rest died a violent death on the battlefield or in palace plots. Considering all 175, 30% were murdered, committed suicide or died in battle.

Researchers at the University of São Paulo’s Institute of Mathematical and Computer Sciences (ICMC-USP) in São Carlos (state of São Paulo, Brazil) investigated the underlying mathematical patterns associated with the reigns of the Roman emperors, showing that they followed what statisticians call a “power law”.

An article on the study is published in Royal Society Open Science, a peer-reviewed scientific journal of the UK’s Royal Society.

“Although they appear to be random, power-law distributions of probabilities are found in many other phenomena associated with complex systems, such as lunar crater sizes, earthquake magnitudes, word frequencies in texts, the market value of companies, and even the number of ‘followers’ people have on social media,” data scientist Francisco Rodrigues, a professor at ICMC-USP and principal investigator for the study, told Agência FAPESP.

The study was funded by FAPESP (projects 19/23293-0 and 17/25971-0), and conducted under the aegis of the Center for Mathematical Sciences Applied to Industry (CeMEAI), a Research, Innovation and Dissemination Center (RIDC) supported by FAPESP and hosted by ICMC-USP.

All the phenomena mentioned by Rodrigues display a pattern often referred to as the Pareto principle or 80/20 rule. Put simply, this means that in all these cases the probability of a common occurrence is about 80% and that of a rare event is about 20%. For example, 80% of lunar craters are relatively small, while 20% are really large. In social media, 80% of users have at most a few dozen followers, while 20% have thousands or even millions. In the case of Roman emperors, the rare event was not being assassinated.

“The first person to observe this ratio was Italian economist Vilfredo Pareto (1848-1923). While studying wealth distribution in Europe, he found that 80% of Italy’s property belonged to 20% of its population. The majority had few resources, and a minority owned most of the wealth,” Rodrigues said.

In addition to the 80/20 rule, another pattern can be seen in the careers of Roman emperors. “When we analyzed time to death for each emperor, we found that the risk was high when the emperor took the throne. This could have something to do with the difficulties and demands of the job and the new emperor’s lack of political expertise. The risk then declines systematically until the emperor has reigned for 13 years. At that point, it rises sharply again,” Rodrigues said.

If the 80/20 rule is a well-known pattern, the sharp downturn in the survival curve around year 13 is a novel finding. “We envisaged several possible explanations for this turning-point. It may be that after the 13-year cycle the emperor’s rivals concluded they were unlikely to ascend the throne by natural means. Perhaps his old enemies regrouped, or new rivals may have come to the fore. A crisis may have arisen owing to all these factors combined. It’s worth noting that the risk falls again after this turning-point,” Rodrigues said.

The change at 13 years is a question that has yet to be answered, but in its pursuit of a long line of quantitative historiography, the paper shows that statistical analysis can be an important complementary resource in the study of historical phenomena. “Historical formations are complex systems in which players interact, collaborate and compete for power and resources. The unpredictable actions of individuals can produce predictable patterns of collective behavior that can be investigated mathematically,” Rodrigues said.

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Ancestral Native American population does not originate in Japan, as believed by many

 A widely accepted theory of Native American origins coming from Japan has been attacked in a new scientific study, which shows that the genetics and skeletal biology "simply does not match-up."

The findings, published today in the peer-reviewed journal PaleoAmerica, are likely to have a major impact on how we understand Indigenous Americans' arrival to the Western Hemisphere.

Based on similarities in stone artifacts, many archaeologists currently believe that Indigenous Americans, or 'First Peoples', migrated to the Americas from Japan about 15,000 years ago.

It is thought they moved along the northern rim of the Pacific Ocean, which included the Bering Land Bridge, until they reached the northwest coast of North America.

From there the First Peoples fanned out across the interior parts of the continent and farther south, reaching the southern tip of South America within less than two thousand years.

The theory is based, in part, on similarities in stone tools made by the 'Jomon' people (an early inhabitant of Japan, 15,000 years ago), and those found in some of the earliest known archaeological sites inhabited by ancient First Peoples.

But this new study, out today in PaleoAmerica -- the flagship journal of the Center for the Study of the First Americans at Texas A&M University -- suggests otherwise.

Carried out by one of the world's foremost experts in the study of human teeth and a team of Ice-Age human genetics experts, the paper analysed the biology and genetic coding of teeth samples from multiple continents and looked directly at the Jomon people.

"We found that the human biology simply doesn't match up with the archaeological theory," states lead author Professor Richard Scott, a recognized expert in the study of human teeth, who led a team of multidisciplinary researchers.

"We do not dispute the idea that ancient Native Americans arrived via the Northwest Pacific coast -- only the theory that they originated with the Jomon people in Japan.

"These people (the Jomon) who lived in Japan 15,000 years ago are an unlikely source for Indigenous Americans. Neither the skeletal biology or the genetics indicate a connection between Japan and the America. The most likely source of the Native American population appears to be Siberia."

In a career spanning almost half a century, Scott -- a professor of anthropology at the University of Nevada-Reno -- has traveled across the globe, collecting an enormous body of information on human teeth worldwide, both ancient and modern. He is the author of numerous scientific papers and several books on the subject.

This latest paper applied multivariate statistical techniques to a large sample of teeth from the Americas, Asia, and the Pacific, showing that quantitative comparison of the teeth reveals little relationship between the Jomon people and Native Americans. In fact, only 7% of the teeth samples were linked to the non-Arctic Native Americans (recognized as the First Peoples).

And, the genetics show the same pattern as the teeth -- little relationship between the Jomon people and Native Americans.

"This is particularly clear in the distribution of maternal and paternal lineages, which do not overlap between the early Jomon and American populations," states co-author Professor Dennis O'Rourke, who was joined by fellow human geneticists -- and expert of the genetics of Indigenous Americans -- at the University of Kansas, Jennifer Raff.

"Plus, recent studies of ancient DNA from Asia reveal that the two peoples split from a common ancestor at a much earlier time," adds Professor O'Rourke.

Together with their colleague and co-author Justin Tackney, O'Rourke and Raff reported the first analysis of ancient DNA from Ice-Age human remains in Alaska in 2016.

Other co-authors include specialists in Ice-Age archaeology and ecology.

Shortly before publication of the paper, two other new studies on related topics were released.

A new genetics paper on the modern Japanese population concluded that it represents three separate migrations into Japan, rather than two, as previously believed. It offered more support to the authors' conclusions, however, about the lack of a biological relationship between the Jomon people and Indigenous Americans.

And, in late September, archaeologists reported in another paper the startling discovery of ancient footprints in New Mexico dating to 23,000 years ago, described as "definitive evidence" of people in North America before the Last Glacial Maximum -- before expanding glaciers probably cut off access from the Bering Land Bridge to the Western Hemisphere. It remains unclear who made the footprints and how they are related to living Native Americans, but the new paper provides no evidence that the latter are derived from Japan.

Professor Scott concludes that "the Incipient Jomon population represents one of the least likely sources for Native American peoples of any of the non-African populations."

Limitations of the study include that available samples of both teeth and ancient DNA for the Jomon population are less than 10,000 years old, i.e., do not antedate the early Holocene (when the First Peoples are understood to arrive in America).

"We assume," the authors explain however, "that they are valid proxies for the Incipient Jomon population or the people who made stemmed points in Japan 16,000-15,000 years ago."

Ancient poop shows people in present-day Austria drank beer and ate blue cheese up to 2,700 years ago

 

Peer-Reviewed Publication

 

IMAGE: THIS IMAGE SHOWS PALEOFECES SAMPLES FROM HALLSTATT SALT MINES ANALYZED IN THIS STUDY. view more 

CREDIT: EURAC RESEARCH/FRANK MAIXNER

Human feces don’t usually stick around for long—and certainly not for thousands of years. But exceptions to this general rule are found in a few places in the world, including prehistoric salt mines of the Austrian UNESCO World Heritage area Hallstatt-Dachstein/Salzkammergut. Now, researchers who’ve studied ancient fecal samples (or paleofeces) from these mines have uncovered some surprising evidence: the presence of two fungal species used in the production of blue cheese and beer. The findings appear in the journal Current Biology on October 13.

“Genome-wide analysis indicates that both fungi were involved in food fermentation and provide the first molecular evidence for blue cheese and beer consumption during Iron Age Europe,” says Frank Maixner (@FrankMaixner) of the Eurac Research Institute for Mummy Studies in Bolzano, Italy.

“These results shed substantial new light on the life of the prehistoric salt miners in Hallstatt and allow an understanding of ancient culinary practices in general on a whole new level,” adds Kerstin Kowarik (@KowarikKerstin) of the Museum of Natural History Vienna. “It is becoming increasingly clear that not only were prehistoric culinary practices sophisticated, but also that complex processed foodstuffs as well as the technique of fermentation have held a prominent role in our early food history.”

Earlier studies already had shown the potential for studies of prehistoric paleofeces from salt mines to offer important insights into early human diet and health. In the new study, Maixner, Kowarik, and their colleagues added in-depth microscopic, metagenomic, and proteomic analyses—to explore the microbes, DNA, and proteins that were present in those poop samples.

These comprehensive studies allowed them to reconstruct the diet of the people who once lived there. They also could get information about the ancient microbes that inhabited their guts. Gut microbes are collectively known as the gut microbiome and are now recognized to have an important role in human health.

Their dietary survey identified bran and glumes of different cereals as one of the most prevalent plant fragments. They report that this highly fibrous, carbohydrate-rich diet was supplemented with proteins from broad beans and occasionally with fruits, nuts, or animal food products.

In keeping with their plant-heavy diet, the ancient miners up to the Baroque period also had gut microbiome structures more like those of modern non-Westernized individuals, whose diets are also mainly composed of unprocessed food, fresh fruits and vegetables. The findings suggest a more recent shift in the Western gut microbiome as eating habits and lifestyles changed.

When the researchers extended their microbial survey to include fungi, that’s when they got their biggest surprise: an abundance in one of their Iron Age samples of Penicillium roquefortiand Saccharomyces cerevisiae DNA.

“The Hallstatt miners seem to have intentionally applied food fermentation technologies with microorganisms which are still nowadays used in the food industry,” Maixner says.

The findings offer the first evidence that people were already producing blue cheese in Iron Age Europe nearly 2,700 years ago, he adds. In ongoing and future studies of the paleofeces from Hallstatt, they hope to learn more about the early production of fermented foods and the interplay between nutrition and the gut microbiome composition in different time periods.

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Marble quarries discovered as source for archaic Apollo

Reports and Proceedings

The source of marble for a statue of Apollo on the Greek island of Delos has been a mystery to art historians and archaeologists for decades. The stone’s chemistry pointed geochemists to the southern end of the nearby island of Naxos, but no one thought there were ancient marble quarries there. A geoarchaeologist believes he found the source.

“We had actually been told that we were not going to find what we were looking for,” says geoarchaeologist Scott Pike of Willamette University. But after two field seasons traipsing across Mediterranean shrublands, Pike believes he has found the source. He is presenting his findings on Monday, 11 October 2021 at the Geological Society of America’s GSA Connects 2021 annual meeting in Portland, Ore.

The Greek Archaic period (approximately 800 to 480 B.C.E.) is known in part for its “larger-than-life” kouros statues, which depicted young men. Together, the massive Apollo kouros on Delos would stand around ten meters (33 feet) high, although today it is broken into several parts. The massive marble chunks are white and worn; at a glance, some of the pieces hardly resemble parts of a human figure. But the statue has drawn researchers all the same. Searching for its source was sparked in part by an ambiguous inscription at its base, roughly translated as, “I am of the same stone, statue and plinth,” with a later addition stating that the kouros was “from the Naxians, to Apollo,” according to Pike.

It was not clear whether the inscription referred to the statue’s structure, being hewn from a single piece of marble, or the origin of its stone. Pike sampled various parts from the statue — a hand, the upper and lower torso, a bit of leg — and analyzed its carbon and oxygen isotopic composition. That composition can be used to trace the marble source by comparing it against other analyzed marbles, like finding a fingerprint match in a database.

“The analyses showed the marble came from Naxos, but from a region where there hasn’t been any evidence of ancient quarrying. We know that there are two quarries in the northern part of the island, where there are still large kouroi in place in the quarries. But we didn’t know of any ancient quarries in the south,” says Pike.

Pike headed to the southern side of Naxos, despite locals assuring him his efforts were in vain. He relied on local knowledge of other archaeological sites and geologic maps to guide him as he “scoured the landscape” looking for, essentially, outcrops of white marble and possible small pits. Remnants of Archaic quarries bear little resemblance to the vast open-pit mines humans create today and were difficult for Pike to find.

After a couple of weeks of searching, Pike began finding small bands of white marble that were not marked on the geologic maps. Some were close to archaeological sites, giving Pike some confidence that these small quarries could be the source.

“Finding what we were looking for was exciting because being told several times that you’re not going to find anything is discouraging, but we knew,” Pike says. “The evidence pointed to the south. I felt the most Indiana Jones I’ll ever be.”

Back in the lab, Pike analyzed his marble samples and found that two of the newly-uncovered southern white marbles were good matches for the Apollo kouros at Delos. Knowing that these early marble quarries exist in the south of the island will be helpful for tracing the source for other ancient marble artifacts, such as older Bronze Age Cycladic figurines that have puzzled geoarchaeologists. It also has implications for knowledge of commerce at the time.

“Knowing now that there is a marble source on Naxos for these Bronze Age statues and figurines will place the region more in the center of commerce, trade and influence than had been previously understood,” Pike says.

 

Finding Apollo’s Marbles: Rediscovery of A White Marble Source in Southern Naxos, Greece
https://gsa.confex.com/gsa/2021AM/meetingapp.cgi/Paper/371225